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  <last_published>2011-11-28T14:46:39</last_published>
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&lt;h3&gt;Per Capita Rates&lt;/h3&gt;
&lt;p&gt;It is important to relate the basic population parameters (such as births or deaths) to the size of the whole population. This allows us to make a better decision if a population is at risk.&lt;/p&gt;
&lt;p&gt; &lt;/p&gt;
&lt;p&gt;We define the &lt;em&gt;per capita birth rate&lt;/em&gt; (or nativity rate) as the number of births per individual per unit time interval: $b=\frac {B}{N}$ .&lt;/p&gt;
&lt;p&gt;Similarly we define the &lt;em&gt;per capita death rate&lt;/em&gt; (or mortality rate) as the number of deaths per individual per unit time interval:  $d=\frac {D}{N}$&lt;/p&gt;
&lt;h3&gt; &lt;/h3&gt;
&lt;h3&gt;The First Model&lt;/h3&gt;
&lt;p&gt;Recall the population equation from &lt;a href=&quot;http://nrich.maths.org/7252?part=index&quot;&gt;before&lt;/a&gt;: $$N_{t+1}=N_t+B-D$$ Because per capita birth and death rates do not change with the size (or density) of the population, we can rewrite our model in terms of per capita rates: $$N_{t+1}=N_t+bN_t-dN_t=N_t+(b-d)N_t$$ This model is said to be &lt;em&gt;density-independent.&lt;/em&gt;&lt;/p&gt;
&lt;p&gt; &lt;/p&gt;
&lt;p&gt;We call the term $r=b-d$, the &lt;em&gt;geometric rate of increase&lt;/em&gt;. Note that $r=\frac{\Delta N_t}{N_t}$ , so &lt;em&gt;r&lt;/em&gt; can be interpreted as the per capita rate of change of population size.&lt;/p&gt;
&lt;p&gt; &lt;/p&gt;
&lt;p&gt;The equation for our model becomes: $$\begin{align*} N_{t+1}&amp;amp;=N_t+rN_t \\ &amp;amp;=(1+r)N_t \\ &amp;amp;=\lambda N_t \end{align*}$$ where $\lambda=1+r$ is defined as the &lt;em&gt;finite rate of increase&lt;/em&gt;. Note that $\lambda=\frac{N_{t+1}}{N_t}$ , so $\lambda$ can be interpreted as the ratio between the population size at one time to another time.&lt;/p&gt;
&lt;h3&gt; &lt;/h3&gt;
&lt;p&gt;How do you think we can solve this new equation? Go &lt;a href=&quot;http://nrich.maths.org/7104?part=index&quot;&gt;here&lt;/a&gt; for more information.&lt;/p&gt;
&lt;p&gt;Do you think this model is valid in reality? What problems do you think might occur? Think about environmental resources and density-independence. An investigation of these problems can be found &lt;a href=&quot;http://nrich.maths.org/7048?part=index&quot;&gt;here&lt;/a&gt;.&lt;/p&gt;
&lt;p&gt; &lt;/p&gt;
&lt;p&gt;&lt;strong&gt;Question:&lt;/strong&gt;&lt;/p&gt;
&lt;p&gt;If 20 sea otters from a total population of 850 are fatally affected by disease, what is the mortality as a per capita rate?&lt;/p&gt;
&lt;p style=&quot;text-align: center;&quot;&gt;&lt;mdo:image style=&quot;width: 300px; height: 200px;&quot; src=&quot;seaotters1-300x200.jpg&quot; alt=&quot;&quot;&gt;&lt;/mdo:image&gt;&lt;/p&gt;
&lt;p&gt;Given the population is initially 850, and increases to 1000 after one year, what is the value of $\lambda$?&lt;/p&gt;
&lt;p&gt;Use this to find the per capita birth rate, and find the population size in ten years.&lt;/p&gt;
&lt;br&gt;&lt;/br&gt;&lt;/mdoxml&gt;</indexXML>
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  <end_user_role>2</end_user_role>
  <difficulty>4</difficulty>
  <keystage1>0</keystage1>
  <keystage2>0</keystage2>
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  <title>Population Dynamics - part 1</title>
  <description>First in our series of problems on population dynamics for advanced students.</description>
  <spec_group>Using, Applying and Reasoning about Mathematics
    <specifier>Mathematical modelling</specifier>
  </spec_group>
  <spec_group>Collections
    <specifier>Population Dynamics</specifier>
  </spec_group>
  <spec_group>Applications
    <specifier>biology</specifier>
  </spec_group>
  <spec_group>Pre-Calculus and Calculus
    <specifier>Calculus generally</specifier>
  </spec_group>
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